Willow doesn't wait till her pups fuss to feed them. She anticipates and wames them up to feed. I believew she is listening to her body. I have milk and I am filling up - it is time. They sometimes fuss when she wakes them up but then they go - oh it is food time.
All pups continue to gain weight. They sleep decently between feedings. One or 2 at a time crawl up into my lap to cuddle. Moms need balance too.
Some videos are up. I will add a few more.